Each level is tuned so that certain advanced techniques have to be used in order to achieve the higher scores, with correspondingly higher grades given to mark progress. Ultimately the game is a high-score attack. The game can be played in a 2-player co-operative mode offering more ways to build and manipulate cell structures or to distract and defeat the Nucleus enemies. There are survival levels where the player is given a source of cells and have to keep back a more and more quickly expanding herd of bacteria and virus by arranging the cells in a free-form way. Other styles of level are also offered-some levels require the player to move cells from one section of the level to another to complete them, some are straight shootouts with no Nucleus enemies, or collect'em-ups in a dynamic environment. There is a sense of zombie pressure as the number of enemies build up and the player tries to keep their defences intact while harvesting protein fast enough.
Exploding protein bombs or angry Nucleus enemies exert a large force over a wide area, which can break the various arrangements of cells the player may have constructed to fend off bacteria or virus. Cells exert an attractive force on each other, clumping into a tesselating mass and reacting to the various forces in the game as a flexing and spongy whole. The game entities are all physically dynamic, each imparting a force on other nearby entities. As the Nucleus enemies get damaged they become more and more violent, producing more spore and eventually attacking the player directly. The player has to attempt to maintain a cell barrier between itself and the possibly multiple, possibly mobile Nucleus enemies, while harvesting as much protein as possible in order to fight them back. These boss-type enemies fire spores at cells to turn them into bacteria, and can only be damaged by protein bombs. Interspersed along the game progression are encounters with super enemies (the Nucleus of the game title). By clumping cells together the player can harvest more protein, creating a trade off between using the cells available to protect their ship and shooting them to release enough protein to be able to fire the more powerful protein bomb weapon. The main resource available to the player is protein, these are released each time an enemy or a cell is destroyed. Each level also has a number of cells which clump together and interact in a physical manner the players can move them around and manipulate them with a traction beam to construct barriers to hide behind or to block of parts of the dynamically changing levels. The basic game consists of shooting bacteria or progressively larger and larger viruses, which break down into smaller viruses when shot, or if left alone combine themselves into ever larger viruses. Players are tasked with ridding a host body of a virus infestation, moving through the body's three main biological systems, the digestive system, the circulatory system, and finally the nervous system, culminating in a final battle in the brain. Nucleus is a twin-stick shooter with puzzle elements for the PlayStation 3 available through the PlayStation Store.
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